The horse (Equus ferus caballus ) is a domesticated, odd-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of Equus ferus. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, Eohippus, into the large, single-toed animal of today. Humans began domesticating horses around 4000 BC, and their domestication is believed to have been widespread by 3000 BC. Horses in the subspecies caballus are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior.
Horses are adapted to run, allowing them to quickly escape predators, possessing an excellent sense of balance and a strong fight-or-flight response. Related to this need to flee from predators in the wild is an unusual trait: horses are able to sleep both standing up and lying down, with younger horses tending to sleep significantly more than adults. Female horses, called mares, carry their young for approximately 11 months, and a young horse, called a foal, can stand and run shortly following birth. Most domesticated horses begin training under a saddle or in a harness between the ages of two and four. They reach full adult development by age five, and have an average lifespan of between 25 and 30 years.
Horse breeds are loosely divided into three categories based on general temperament: spirited "hot bloods" with speed and endurance; "cold bloods", such as draft horses and some ponies, suitable for slow, heavy work; and "warmbloods", developed from crosses between hot bloods and cold bloods, often focusing on creating breeds for specific riding purposes, particularly in Europe. There are more than 300 breeds of horse in the world today, developed for many different uses.
Horses and humans interact in a wide variety of sport competitions and non-competitive recreational pursuits, as well as in working activities such as police work, agriculture, entertainment, and therapy. Horses were historically used in warfare, from which a wide variety of riding and driving techniques developed, using many different styles of equipment and methods of control. Many products are derived from horses, including meat, milk, hide, hair, bone, and pharmaceuticals extracted from the urine of pregnant mares. Humans provide domesticated horses with food, water, and shelter, as well as attention from specialists such as veterinarians and farriers.
Modern horses are often used to reenact many of their historical work purposes. Horses are used, complete with equipment that is authentic or a meticulously recreated replica, in various live action historical reenactments of specific periods of history, especially recreations of famous battles. Horses are also used to preserve cultural traditions and for ceremonial purposes. Countries such as the United Kingdom still use horse-drawn carriages to convey royalty and other VIPs to and from certain culturally significant events. Public exhibitions are another example, such as the Budweiser Clydesdales, seen in parades and other public settings, a team of draft horses that pull a beer wagon similar to that used before the invention of the modern motorized truck.
Horses are frequently used in television, films and literature. They are sometimes featured as a major character in films about particular animals, but also used as visual elements that assure the accuracy of historical stories. Both live horses and iconic images of horses are used in advertising to promote a variety of products. The horse frequently appears in coats of arms in heraldry, in a variety of poses and equipment. The mythologies of many cultures, including Greco-Roman, Hindu, Islamic, and Norse, include references to both normal horses and those with wings or additional limbs, and multiple myths also call upon the horse to draw the chariots of the Moon and Sun. The horse also appears in the 12-year cycle of animals in the Chinese zodiac related to the Chinese calendar. Horses serve as the inspiration for many modern automobile names and logos, including the Ford Pinto, Ford Bronco, Ford Mustang, Hyundai Equus, Hyundai Pony, Mitsubishi Starion, Subaru Brumby, Mitsubishi Colt/Dodge Colt, Volkswagen Polo, Pinzgauer, Steyr-Puch Haflinger, Pegaso, Porsche, Rolls-Royce Camargue, Ferrari, Carlsson, Kamaz, Corre La Licorne, Iran Khodro, Eicher, and Baojun.
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ViviparousAmong animals, viviparity is the development of the embryo inside the body of the parent. The term 'viviparity' and its adjective form 'viviparous'...
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PrecocialPrecocial species are those in which the young are relatively mature and mobile from the moment of birth or hatching. Precocial species are normall...
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HerdingA herd is a social grouping of certain animals of the same species, either wild or domestic. The form of collective animal behavior associated with...
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Dominance hierarchyA dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social gr...
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DomesticatedDomesticated animals are those adapted to live with humans. It is the mutual relationship between animals and humans who have an influence on their...
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starts withThe height of horses is measured at the highest point of the withers, where the neck meets the back. This point is used because it is a stable point of the anatomy, unlike the head or neck, which move up and down in relation to the body of the horse.
In English-speaking countries, the height of horses is often stated in units of hands and inches: one hand is equal to 4 inches (101.6 mm). The height is expressed as the number of full hands, followed by a point, then the number of additional inches, and ending with the abbreviation "h" or "hh" (for "hands high"). Thus, a horse described as "15.2 h" is 15 hands plus 2 inches, for a total of 62 inches (157.5 cm) in height.
The size of horses varies by breed, but also is influenced by nutrition. Light riding horses usually range in height from 14 to 16 hands (56 to 64 inches, 142 to 163 cm) and can weigh from 380 to 550 kilograms (840 to 1,210 lb). Larger riding horses usually start at about 15.2 hands (62 inches, 157 cm) and often are as tall as 17 hands (68 inches, 173 cm), weighing from 500 to 600 kilograms (1,100 to 1,320 lb). Heavy or draft horses are usually at least 16 hands (64 inches, 163 cm) high and can be as tall as 18 hands (72 inches, 183 cm) high. They can weigh from about 700 to 1,000 kilograms (1,540 to 2,200 lb).
The largest horse in recorded history was probably a Shire horse named Mammoth, who was born in 1848. He stood 21.2 1⁄4 hands (86.25 inches, 219 cm) high and his peak weight was estimated at 1,524 kilograms (3,360 lb). The record holder for the smallest horse ever is Thumbelina, a fully mature miniature horse affected by dwarfism. She was 17 in (43 cm) tall and weighed 57 lb (26 kg).
Horses exhibit a diverse array of coat colors and distinctive markings, described by a specialized vocabulary. Often, a horse is classified first by its coat color, before breed or sex. Horses of the same color may be distinguished from one another by white markings, which, along with various spotting patterns, are inherited separately from coat color.
Many genes that create horse coat colors and patterns have been identified. Current genetic tests can identify at least 13 different alleles influencing coat color, and research continues to discover new genes linked to specific traits. The basic coat colors of chestnut and black are determined by the gene controlled by the Melanocortin 1 receptor, also known as the "extension gene" or "red factor," as its recessive form is "red" (chestnut) and its dominant form is black. Additional genes control suppression of black color to point coloration that results in a bay, spotting patterns such as pinto or leopard, dilution genes such as palomino or dun, as well as graying, and all the other factors that create the many possible coat colors found in horses.
Horses that have a white coat color are often mislabeled; a horse that looks "white" is usually a middle-aged or older gray. Grays are born a darker shade, get lighter as they age, but usually keep black skin underneath their white hair coat (with the exception of pink skin under white markings). The only horses properly called white are born with a predominantly white hair coat and pink skin, a fairly rare occurrence. Different and unrelated genetic factors can produce white coat colors in horses, including several different alleles of dominant white and the sabino-1 gene. However, there are no "albino" horses, defined as having both pink skin and red eyes.
All horses move naturally with four basic gaits:
natomy, different life stages, and colors and breeds.
All horses move naturally with four basic gaits:
Besides these basic gaits, some horses perform a two-beat pace, instead of the trot. There also are several four-beat 'ambling' gaits that are approximately the speed of a trot or pace, though smoother to ride. These include the lateral rack, running walk, and tölt as well as the diagonal fox trot. Ambling gaits are often genetic in some breeds, known collectively as gaited horses. These horses replace the trot with one of the ambling gaits.
Horses are prey animals with a strong fight-or-flight response. Their first reaction to a threat is to startle and usually flee, although they will stand their ground and defend themselves when flight is impossible or if their young are threatened. They also tend to be curious; when startled, they will often hesitate an instant to ascertain the cause of their fright, and may not always flee from something that they perceive as non-threatening. Most light horse riding breeds were developed for speed, agility, alertness and endurance; natural qualities that extend from their wild ancestors. However, through selective breeding, some breeds of horses are quite docile, particularly certain draft horses.
Horses are herd animals, with a clear hierarchy of rank, led by a dominant individual, usually a mare. They are also social creatures that are able to form companionship attachments to their own species and to other animals, including humans. They communicate in various ways, including vocalizations such as nickering or whinnying, mutual grooming, and body language. Many horses will become difficult to manage if they are isolated, but with training, horses can learn to accept a human as a companion, and thus be comfortable away from other horses. However, when confined with insufficient companionship, exercise, or stimulation, individuals may develop stable vices, an assortment of bad habits, mostly stereotypies of psychological origin, that include wood chewing, wall kicking, "weaving" (rocking back and forth), and other problems.
Gestation lasts approximately 340 days, with an average range 320–370 days, and usually results in one foal; twins are rare. Horses are a precocial species, and foals are capable of standing and running within a short time following birth. Foals are usually born in the spring. The estrous cycle of a mare occurs roughly every 19–22 days and occurs from early spring into autumn. Most mares enter an anestrus period during the winter and thus do not cycle in this period. Foals are generally weaned from their mothers between four and six months of age.
Horses, particularly colts, are sometimes physically capable of reproduction at about 18 months, but domesticated horses are rarely allowed to breed before the age of three, especially females. Horses four years old are considered mature, although the skeleton normally continues to develop until the age of six; maturation also depends on the horse's size, breed, sex, and quality of care. Larger horses have larger bones; therefore, not only do the bones take longer to form bone tissue, but the epiphyseal plates are larger and take longer to convert from cartilage to bone. These plates convert after the other parts of the bones, and are crucial to development.
Depending on maturity, breed, and work expected, horses are usually put under saddle and trained to be ridden between the ages of two and four. Although Thoroughbred race horses are put on the track as young as the age of two in some countries, horses specifically bred for sports such as dressage are generally not put under saddle until they are three or four years old, because their bones and muscles are not solidly developed. For endurance riding competition, horses are not deemed mature enough to compete until they are a full 60 calendar months (five years) old.
Domestication of the horse most likely took place in central Asia prior to 3500 BC. Two major sources of information are used to determine where and when the horse was first domesticated and how the domesticated horse spread around the world. The first source is based on palaeological and archaeological discoveries; the second source is a comparison of DNA obtained from modern horses to that from bones and teeth of ancient horse remains.
The earliest archaeological evidence for the domestication of the horse comes from sites in Ukraine and Kazakhstan, dating to approximately 4000–3500 BC. By 3000 BC, the horse was completely domesticated and by 2000 BC there was a sharp increase in the number of horse bones found in human settlements in northwestern Europe, indicating the spread of domesticated horses throughout the continent. The most recent, but most irrefutable evidence of domestication comes from sites where horse remains were interred with chariots in graves of the Sintashta and Petrovka cultures c. 2100 BC.
A 2021 genetic study suggested that most modern domestic horses descend from the lower Volga-Don region. Ancient horse genomes indicate that these populations influenced almost all local populations as they expanded rapidly throughout Eurasia, beginning about 4200 years ago. It also shows that certain adaptations were strongly selected due to riding, and that equestrian material culture, including Sintashta spoke-wheeled chariots spread with the horse itself.
Domestication is also studied by using the genetic material of present-day horses and comparing it with the genetic material present in the bones and teeth of horse remains found in archaeological and palaeological excavations. The variation in the genetic material shows that very few wild stallions contributed to the domestic horse, while many mares were part of early domesticated herds. This is reflected in the difference in genetic variation between the DNA that is passed on along the paternal, or sire line (Y-chromosome) versus that passed on along the maternal, or dam line (mitochondrial DNA). There are very low levels of Y-chromosome variability, but a great deal of genetic variation in mitochondrial DNA. There is also regional variation in mitochondrial DNA due to the inclusion of wild mares in domestic herds. Another characteristic of domestication is an increase in coat color variation. In horses, this increased dramatically between 5000 and 3000 BC.
Before the availability of DNA techniques to resolve the questions related to the domestication of the horse, various hypotheses were proposed. One classification was based on body types and conformation, suggesting the presence of four basic prototypes that had adapted to their environment prior to domestication. Another hypothesis held that the four prototypes originated from a single wild species and that all different body types were entirely a result of selective breeding after domestication. However, the lack of a detectable substructure in the horse has resulted in a rejection of both hypotheses.
Social animals are those animals that interact highly with other animals, usually of their own species (conspecifics), to the point of having a rec...