The analysis of morphological, behavioural, and especially genetic characters have since identified further cryptic species of whiskered bats in the genus Myotis, including Myotis alcathoe (described in 2001 from Europe). Myotis aurascens and Myotis ikonnikovi are other similar species. Myotis hajastanicus was also included in M. mystacinus until recently, but it was differentiated on the base of morphologic comparison.